1. Biotechnology Research Center, Southwest University , Chongqing 400716 , China
2. School of Life Sciences, Chongqing University , Chongqing 401331 , China
3. School of Agricultural Sciences, Zhengzhou University , Zhengzhou, Henan 450001 , China
4. College of Agronomy and Biotechnology, Southwest University , Chongqing 400716 , China