1. Department of Animal and Plant Biology, State University of Londrina (UEL) , Londrina, PR, 86057-970 , Brazil
2. Center for Natural and Human Sciences, Federal University of ABC (UFABC) , Santo André, SP 09210-580 , Brazil
3. Department of Plant Biology, University of Szeged , Szeged, 6726 , Hungary
4. Doctoral School of Biology, Faculty of Science and Informatics, University of Szeged , Szeged, 6726 , Hungary