1. Graduate School of Life Sciences, Tohoku University , 2-1-1 Katahira, Aoba-ku, Sendai 980 - 8577 , Japan
2. Graduate School of Science and Engineering, Yamagata University , 1-4-12 Kojirakawa-machi, Yamagata 990-8560 , Japan
3. Faculty of Science, Yamagata University , 1-4-12 Kojirakawa-machi, Yamagata 990-8560 , Japan
4. Centre for Plant Integrative Biology, University of Nottingham , Nottingham LE12 5RD , UK
5. Plant and Crop Sciences, School of Biosciences, University of Nottingham , Nottingham LE12 5RD , UK
6. Research Center for Space Agriculture and Horticulture, Graduate School of Horticulture, Chiba University , 648 Matsudo, Chiba 271-8510 , Japan