1. Department of Plant Science, The Pennsylvania State University , University Park , PA 16802 , USA
2. Department of Ecosystem Science and Management, The Pennsylvania State University , University Park , PA 16802 , USA
3. Graduate Program in Ecology, The Pennsylvania State University , University Park , PA 16802 , USA
4. Department of Plant Pathology and Environmental Microbiology, The Pennsylvania State University , University Park , PA 16802 , USA