1. Jiangsu Key Laboratory of Crop Genetics and Physiology/ Jiangsu Key Laboratory of Crop Cultivation and Physiology, Agricultural College of Yangzhou University , Yangzhou 225009 , China
2. Jiangsu Co-Innovation Center for Modern Production Technology of Grain Crops, Yangzhou University , Yangzhou 225009 , China
3. Department of Biology, Hong Kong Baptist University , Hong Kong 999077 , China