1. Graduate School of Science and Technology, Nara Institute of Science and Technology , 8916-5 Takayama, Ikoma, Nara 630-0192 , Japan
2. School of Biological Science and Technology, College of Science and Engineering, Kanazawa University , Kakuma-machi, Kanazawa, Ishikawa 920-1192 , Japan
3. RIKEN Center for Sustainable Resource Science , Tsurumi-ku, Yokohama, Kanagawa 230-0045 , Japan
4. Department of Biological Sciences, The University of Tokyo , Tokyo, 119-0033 , Japan
5. RIKEN Cluster for Pioneering Research , 2-1 Hirosawa, Wako, Saitama 351-0198 , Japan