1. the Chinese Academy of Sciences Key Laboratory of Plant Resources, Institute of Botany, , Beijing 100093, China
2. China National Botanical Garden , Beijing 100093, China
3. College of Life Science, University of Chinese Academy of Sciences , Beijing 100049, China
4. Qingdao Agricultural University College of Landscape Architecture and Forestry, , Qingdao 266109, China
5. Peony Research Institute, Luoyang Academy of Agricultural and Forestry Sciences , Luoyang 471000, China