1. Department of Molecular Physiology and Biophysics,
2. Stephen A. Wynn Institute for Vision Research, University of Iowa, Iowa City, IA 52242, USA
3. Department of Ophthalmology and Visual Sciences, Carver College of Medicine, University of Iowa, Iowa City, IA 52242, USA
4. Center for Prevention and Treatment of Visual Loss, Iowa City Veterans Administration Medical Center, Iowa City, IA 52242, USA