1. Department of Biology, Lehrstuhl für Zoologie und Evolutionsbiologie, University of Konstanz, Konstanz, Germany
2. Present address: Molecular Ecology and Evolution Laboratory, Department of Biological Science, Sangji University, Wonju, Korea
3. Present address: Hellenic Centre for Marine Research (HCMR), Institute of Marine Biology, Biotechnology, and Aquaculture (IMBBC), Heraklion, Greece
4. Present address: Korean Entomological Institute, Korea University, Seoul, Korea
5. Present address: Department of Collective Behaviour, Max Planck Institute for Ornithology and University of Konstanz, Konstanz, Germany