1. Centre for Genetics and Genomics Versus Arthritis, Centre for Musculoskeletal Research, The University of Manchester , Manchester, UK
2. NIHR Biomedical Research Centre Manchester, Manchester University NHS Foundation Trust, Manchester Academic Health Science Centre , Manchester, UK
3. School of Medicine, University of Leeds , Leeds, UK
4. NIHR Leeds Biomedical Research Centre, Leeds Teaching Hospitals NHS Trust , Leeds, UK
5. NIHR In Vitro Diagnostic Co-operative, Leeds Teaching Hospitals NHS Trust , Leeds, UK
6. Translational and Clinical Research Institute, Newcastle University , Newcastle-upon-Tyne, UK
7. Musculoskeletal Unit, Newcastle-upon-Tyne Hospitals NHS Foundation Trust , Newcastle-upon-Tyne, UK
8. School of Medicine and Medical Science, Conway Institute, University College Dublin , Dublin, Ireland
9. Centre for Epidemiology Versus Arthritis, Centre for Musculoskeletal Research, The University of Manchester , Manchester, UK