1. Versus Arthritis Centre for Genetics and Genomics, Division of Musculoskeletal Sciences, The University of Manchester
2. NIHR Manchester Biomedical Research Centre, Manchester University NHS Foundation Trust, Manchester Academic Health Sciences Centre , Manchester
3. School of Medicine, University of Leeds & NIHR Leeds Biomedical Research Centre and NIHR In Vitro Diagnostic Co-operative, Leeds Teaching Hospitals NHS Trust, University of Leeds , Leeds
4. Translational & Clinical Research Institution, Newcastle University & Musculoskeletal Unit, Newcastle upon Tyne Hospitals NHS Foundation Trust, Newcastle University , Newcastle upon Tyne, UK
5. School of Medicine & Medical Science, Conway Institute, University College Dublin , Bellfield, Dublin 4, Ireland
6. Versus Arthritis Centre for Epidemiology, Centre for Musculoskeletal Research
7. Lydia Becker Institute of Immunology and Inflammation, Faculty of Biology, Medicine and Health, The University of Manchester , Manchester, UK