1. Institute of Life Course and Medical Sciences, University of Liverpool , Liverpool
2. Department of Rheumatology, Countess of Chester Hospital NHS Foundation Trust , Chester, UK
3. Department of Genetics, Cell- and Immunobiology
4. Department of Rheumatology and Clinical Immunology, Department of Internal Medicine and Oncology, Semmelweis University, Budapest, Hungary
5. Centre for Rheumatic Diseases, King's College London , London, UK