1. Group for Mucosal Immunity and Pathogen Agents (GIMAP), EA 3064, University of Lyon, St-Etienne, France
2. Laboratory of Infectious Agents and Hygiene, University Hospital of St-Etienne, St-Etienne cedex 02, France
3. Department of Ear Nose Throat Surgery, University Hospital of St-Etienne, St-Etienne cedex 02, France
4. Corneal Graft Biology, Engineering and Imaging Laboratory (BiiGC), EA2521, University of Lyon, St-Etienne, France
5. Department of Infectious Diseases, University Hospital of St-Etienne, St-Etienne cedex 02, France