1. Institute of Veterinary Science, University of Liverpool, Leahurst Campus, Neston, UK
2. Department of Epidemiology and Population Health, Institute of Infection and Global Health, University of Liverpool, Leahurst Campus, Neston, UK
3. NIHR Health Protection Research Unit in Emerging and Zoonotic Infections, Liverpool, UK
4. The Royal (Dick) School of Veterinary Studies, Easter Bush Campus, Midlothian, UK