1. Department of Integrative Biology, University of California, Berkeley, CA 94720, USA
2. Channel Islands National Park, Ventura, CA 93001, USA
3. Santa Barbara Botanic Garden, Santa Barbara, CA 93105, USA
4. The Marmot Society, South Lake Tahoe, CA 96150, USA
5. Center for Ecosystem Science and Society, Northern Arizona University, Flagstaff, AZ 86011, USA