1. Forestry and Forest Products Research Institute Department of Plant Ecology, , 1 Matsunosato, Tsukuba, Ibaraki 305-8687 , Japan
2. Forestry and Forest Products Research Institute Shikoku Research Center, , 2-915, Asakuranishi, Kochi 780-8077 , Japan
3. Kyushu University Department of Biology, Faculty of Science, , 744, Motooka, Fukuoka 819-0395 , Japan