1. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Menglun, Mengla 666303, Yunnan, China
2. Center of Plant Ecology, Core Botanical Gardens, Chinese Academy of Sciences, Menglun, Mengla 666303, Yunnan, China
3. University of Chinese Academy of Sciences, Yuquan Road 19A, Beijing 100049, China
4. Hawkesbury Institute for the Environment, Western Sydney University, Locked Bag 1797, Penrith NSW 2751, Australia
5. Lijiang Forest Ecosystem Research Station, Kunming Institute of Botany, Chinese Academy of Sciences, Lijiang 674100, Yunnan, China
6. Department of Renewable Resources, University of Alberta, Edmonton, Alberta T6G 2H1, Canada