1. Department of Environmental and Biological Sciences, University of Eastern Finland, Yliopistokatu 7, PO Box 111, 80101 Joensuu, Finland
2. Organismal and Evolutionary Biology Research Programme, University of Helsinki, Viikinkaari 1, PO Box 65, 00014 Helsinki, Finland
3. School of Biological Sciences, Nanyang Technological University, 60 Nanyang Drive, Singapore 637551, Singapore