1. Guangdong Laboratory for Lingnan Modern Agriculture, South China Agricultural University , Wushan Road, Guangzhou 510642 , China
2. College of Biology and Food Engineering, Guangdong University of Education , Guangzhou 510303 , China
3. Guangdong Technology Research Center for Traditional Chinese Veterinary Medicine and Natural Medicine, South China Agricultural University , Wushan Road, Guangzhou 510642 , China
4. Guangdong Key Laboratory for Innovative Development and Utilization of Forest Plant Germplasm, South China Agricultural University , Wushan Road, Guangzhou 510642 , China