1. Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110010, China
2. College of Resources and Environment, University of Chinese Academy of Sciences, Beijing 100049, China
3. Department of Biology, University of Miami, PO Box 249118, Coral Gables, FL 33124, USA
4. Departamento de Ecología, Genética y Evolución, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Universitaria, Nuñez, Buenos Aires C1428EGA, Argentina