1. State Key Laboratory of Crop Biology, MOA Key Laboratory of Horticultural Crop Biology and Germplasm Innovation, College of Horticulture Science and Engineering, Shandong Agricultural University, Tai-An, Shandong, China
2. Research Institute of Pomology, Chinese Academy of Agricultural Sciences, Xingcheng, Liaoning, China
3. College of Life Science, Shandong Agricultural University, Tai-An, Shandong, China