1. Biodiversity Division, National Institute for Environmental Studies , 16-2 Onogawa, Tsukuba, Ibaraki 305-8506 , Japan
2. Graduate School of Bioresources, Mie University , 1577 Kurimamachiya, Mie, Tsu 514-8507 , Japan
3. Faculty of Agriculture and Life Science, Hirosaki University , 3 Bunkyo-cho, Hirosaki, Aomori 036-8561 , Japan