1. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems, South China Botanical Garden, Chinese Academy of Sciences, 723 Xingke Road, Tianhe District, Guangzhou 510650, China
2. Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, Chinese Academy of Sciences, 723 Xingke Road, Tianhe District, Guangzhou 510650, China
3. Département des Sciences Fondamentales, Université du Québec à Chicoutimi, 555 Boulevard de l'Universite, QC G7H2B1, Canada
4. University of Chinese Academy of Sciences, 19(A) Yuquan Road, Shijingshan District, Beijing 100049, China