1. College of Biological Sciences and Biotechnology, Beijing Forestry University, No 35, Qinghua East Road, Haidian District, Beijing 100083, People's Republic of China
2. School of Biotechnology, Jiangsu University of Science and Technology, ZhenJiang, Jiangsu 212003, People's Republic of China
3. National Engineering Laboratory for Tree Breeding, the Tree and Ornamental Plant Breeding and Biotechnology Laboratory of Chinese Forestry Administration, Beijing 100083, People’s Republic of China