1. Forest Bio-Research Center, Forestry and Forest Products Research Institute, Hitachi, Ibaraki, Japan
2. Division of Forest and Biomaterials Science, Graduate School of Agriculture, Kyoto University, Kyoto, Japan
3. Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, Ibaraki, Japan
4. Research Faculty of Agriculture, Hokkaido University, Sapporo, Japan
5. Forest Tree Breeding Center, Forestry and Forest Products Research Institute, Hitachi, Ibaraki, Japan