1. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization & Ecological Restoration Biodiversity Conservation Key Laboratory of Sichuan Province, Chengdu Institute of Biology, Chinese Academy of Sciences, PO Box 416, Chengdu 610041, PR China
2. Nicholas School of the Environment, Box 90328, Duke University, Durham, NC 27708, USA
3. Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Menglun, Mengla, Yunnan 666303, China