1. Department of Forestry and Wildland Resources, Humboldt State University, 1 Harpst Street, Arcata, CA 95521, USA
2. Department of Biological Sciences, Humboldt State University, 1 Harpst Street, Arcata, CA 95521, USA
3. Department of Wildlife, Humboldt State University, 1 Harpst Street, Arcata, CA 95521, USA