1. Institute of Horticultural Biotechnology , , No. 15 Shangxiadian Road, Cangshan District, Fuzhou City, Fujian 350002 , China
2. Fujian Agriculture and Forestry University , , No. 15 Shangxiadian Road, Cangshan District, Fuzhou City, Fujian 350002 , China
3. Institut de la Recherche Interdisciplinaire de Toulouse , , 31300 Toulouse , France
4. IRIT-ARI , , 31300 Toulouse , France