1. Organization for Marine Science and Technology, Nagasaki University, 1551-7 Tairamachi, Nagasaki 851-2213, Japan
2. Graduate School of Fisheries and Environmental Sciences, Nagasaki University, 1-14 Bunkyomachi, Nagasaki 851-8521, Japan
3. Department of Biology, Natural Science Division, Pepperdine University, 24255 Pacific Coast Highway, Malibu, CA 90263-4321, USA