1. Fisheries Laboratory, Graduate School of Agricultural and Life Sciences, University of Tokyo, Hamamatsu, Shizuoka, Japan
2. Institute of Molecular and Cell Biology, A*STAR, Biopolis, Singapore, Singapore
3. National Research Institute of Fisheries Science, Fisheries Research Agency, Yokohama, Kanagawa, Japan
4. Laboratory of Bioresources, National Institute for Basic Biology, Okazaki, Aichi, Japan