1. Plant Biology and Crop Science, Rothamsted Research, West Common, Harpenden, Hertfordshire, UK
2. Lancaster Environment Centre, Lancaster University, Bailrigg, Lancaster, UK
3. School of Biological Sciences, University of Essex, Wivenhoe Park, Colchester, Essex, UK
4. Centre for Crop Systems Analysis, Wageningen University, Wageningen AK, The Netherlands
5. Computational and Systems Biology, Rothamsted Research, West Common, Harpenden, Hertfordshire, UK