1. Ministry of Education Key Laboratory of Plant Genetic Improvement and Comprehensive Utilization, Fujian Agriculture and Forestry University, Fuzhou, China
2. Key Laboratory of Applied Genetics of Universities in Fujian Province, Fujian Agriculture and Forestry University, Fuzhou, Fujian, China
3. College of Crop Science, Fujian Agriculture and Forestry University, Fuzhou, China
4. These authors contributed equally to the paper
5. Department of Botany Government College University, Faisalabad, Pakistan