1. School of Mathematics and Statistics, Beijing Jiaotong University , Beijing, 100044 , China
2. Key Laboratory of Computational Biology, Shanghai Institute of Nutrition and Health, University of Chinese Academy of Sciences, Chinese Academy of Sciences , Shanghai 200031 , China
3. State Key Laboratory of Genetic Engineering , Human Phenome Institute, Zhangjiang Fudan International Innovation Center, Center for Evolutionary Biology, School of Life Sciences, Department of Liver Surgery and Transplantation Liver Cancer Institute, Zhongshan Hospital, , Shanghai 200032 , China
4. Fudan University , Human Phenome Institute, Zhangjiang Fudan International Innovation Center, Center for Evolutionary Biology, School of Life Sciences, Department of Liver Surgery and Transplantation Liver Cancer Institute, Zhongshan Hospital, , Shanghai 200032 , China
5. Ministry of Education Key Laboratory of Contemporary Anthropology , Collaborative Innovation Center for Genetics and Development, , Shanghai 201203 , China
6. Fudan University , Collaborative Innovation Center for Genetics and Development, , Shanghai 201203 , China
7. School of Life Science and Technology, ShanghaiTech University , Shanghai 201210 , China