1. Shandong University School of Software, , Jinan 250101 , China
2. Shandong University Joint SDU-NTU Centre for Artificial Intelligence Research (C-FAIR), , Jinan 250101 , China
3. King Abdullah University of Science and Technology (KAUST) Computer Science Program, Computer, Electrical and Mathematical Sciences and Engineering Division (CEMSE), , Thuwal , Saudi Arabia
4. King Abdullah University of Science and Technology (KAUST) Computational Bioscience Research Center (CBRC), , Thuwal , Saudi Arabia
5. The University of Tokyo Department of Computational Biology and Medical Sciences, , 5-1-5 Kashiwanoha, Kashiwa-shi, Chiba 277-8562 , Japan
6. The Institute of Medical Science, The University of Tokyo Human Genome Center, , 4-6-1 Shirokanedai Minato-ku, Tokyo 108-8639 , Japan
7. University of Electronic Science and Technology of China Institute of Fundamental and Frontier Sciences, , Chengdu, 610054 , China