1. State Key Laboratory of Plant Diversity and Specialty Crops/Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations , Kunming Institute of Botany, , Kunming 650201 Yunnan , China
2. Chinese Academy of Sciences , Kunming Institute of Botany, , Kunming 650201 Yunnan , China
3. University of Chinese Academy of Sciences , Beijing 101408 Beijing , China
4. Institute of Crop Germplasm Resources, Shandong Academy of Agricultural Sciences , Jinan 250100 Shandong , China