1. Medical Scientist Training Program, University of Pittsburgh, Pittsburgh, PA 15213, USA
2. Translational Neuroscience Program, Department of Psychiatry, School of Medicine, University of Pittsburgh, Pittsburgh, PA 15213, USA
3. Center for the Neural Basis of Cognition, Carnegie Mellon University, Pittsburgh, PA 15213, USA
4. Department of Neuroscience, Dietrich School of Arts and Sciences, University of Pittsburgh, Pittsburgh, PA 15213, USA