1. Laboratory for Cognitive Neuroscience , Graduate School of Frontier Biosciences, , 1-4 Yamadaoka, Suita, Osaka 565-0871, Japan
2. Osaka University , Graduate School of Frontier Biosciences, , 1-4 Yamadaoka, Suita, Osaka 565-0871, Japan
3. Center for Information and Neural Networks, National Institute of Information and Communications Technology and Osaka University , 1-4 Yamadaoka, Suita, Osaka 565-0871, Japan
4. Systems Neuroscience Section , Primate Research Institute, , 41-2 Kanrin, Inuyama, Aichi 484-8506, Japan
5. Kyoto University , Primate Research Institute, , 41-2 Kanrin, Inuyama, Aichi 484-8506, Japan