1. Department of Molecular Physiology, Kyushu University Graduate School of Medical Sciences, Fukuoka 812-8582, Japan
2. Friedrich Miescher Institute for Biomedical Research, Basel 4058, Switzerland
3. Division of Visual Information Processing, National Institute for Physiological Sciences, National Institutes of Natural Sciences, Okazaki 444-8585, Japan
4. Department of Physiological Sciences, The Graduate University for Advanced Studies, Okazaki 444-8585, Japan
5. Keio University School of Medicine, Shinanomachi, Shinjuku-ku, 160-8582, Japan
6. Department of Biophysics, Kyoto University Graduate School of Science, Kyoto 606-8502, Japan
7. Department of Physiology, Graduate School of Medical and Dental Sciences, Kagoshima University, Kagoshima 890-8544, Japan
8. CREST, Japan Science and Technology Agency, Kawaguchi, Saitama 332-0012, Japan
9. Department of Physiology, The University of Tokyo School of Medicine, Tokyo 113-0033, Japan
10. International Research Center for Neurointelligence (WPI-IRCN), The University of Tokyo School of Medicine, Tokyo 113-0033, Japan
11. Institute for AI and Beyond, The University of Tokyo School of Medicine, Tokyo 113-0033, Japan