1. Cellular and Molecular Biology Program, University of Wisconsin-Madison , Madison, WI 53705 , United States
2. Biophysics Program, University of Wisconsin-Madison , Madison, WI 53705 , United States
3. Waisman Center, University of Wisconsin-Madison , Madison, WI 53705 , United States
4. University of Wisconsin-Madison Department of Neuroscience, , Madison, WI 53705 , United States