1. Department of Neuroscience, Physiology & Pharmacology (NPP), University College London, London WC1E 6BT, UK
2. Present address: MRC Laboratory of Molecular Biology, Francis Crick Avenue, Cambridge Biomedical Campus, Cambridge CB2 0QH, UK
3. Present address: Queensland Brain Institute, The University of Queensland, St Lucia, QLD 4072, Australia
4. Department of Behavioural Science and Health, University College London, London WC1E 6BT, UK
5. The Queen Square Brain Bank for Neurological Disorders, Department of Molecular Neuroscience, UCL Institute of Neurology, London, UK
6. Division of Neuropathology and Department of Neurodegenerative Disease, Institute of Neurology, University College London, London WC1N 3BG, UK