1. S , Shanghai 201210 , China
2. chool of Life Science and Technology, ShanghaiTech University , Shanghai 201210 , China
3. Shanghai-MOST Key Laboratory of Health and Disease Genomics, NHC Key Lab of Reproduction Regulation, Shanghai Institute for Biomedical and Pharmaceutical Technologies , Shanghai 200237 , China
4. Center for Excellence in Molecular Cell Science, Shanghai Institute of Biochemistry and Cell Biology, Chinese Academy of Sciences , Shanghai 200031 , China
5. Department of Biological Sciences, University of Delaware , Newark, DE 19716 , USA