1. The University of Tokyo Laboratory of Chromatin Structure and Function, Institute for Quantitative Biosciences, , 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-0032, Japan
2. The University of Tokyo Department of Biological Sciences, Graduate School of Science, , 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-0032, Japan
3. RIKEN Center for Biosystems Dynamics Research , 1-7-22 Suehiro-cho, Tsurumi-ku, Yokohama 230-0045, Japan