1. Nagasaki University Graduate School of Biomedical Sciences , Nagasaki 852-8523, Japan
2. Nagasaki University School of Medicine Department of Biochemistry, , Nagasaki 852-8523, Japan
3. Nagasaki University Graduate School of Biomedical Sciences Department of Dermatology, , Nagasaki 852-8523, Japan
4. Nagasaki University School of Medicine Biomedical Research Support Center, , Nagasaki 852-8523, Japan
5. Nagasaki University School of Medicine Department of Histology and Cell Biology, , Nagasaki 852-8523, Japan