1. Department of Agricultural, Forest and Food Sciences, University of Torino , Largo P. Braccini 2, 10095 Grugliasco , Italy
2. Department of Veterinary Sciences, University of Torino , Largo P. Braccini 2, 10095 Grugliasco , Italy
3. Wageningen University & Research, Wageningen Livestock Research , De Elst 1, 6700 AH Wageningen , the Netherlands