1. Department of Food and Life Science, Obihiro University of Agriculture and Veterinary Medicine , Obihiro, Hokkaido 080-8555 , Japan
2. Graduate School of Biostudies, Kyoto University , Kyoto 606-8502 , Japan
3. Research Center for Global Agromedicine, Obihiro University of Agriculture and Veterinary Medicine , Obihiro, Hokkaido 080-8555 , Japan