1. School of Life Sciences, University of KwaZulu-Natal , Pietermaritzburg , South Africa
2. Okavango Research Institute, University of Botswana , Maun , Botswana
3. Agricultural Research Service, United States Department of Agriculture , Miles City, Montana , USA
4. Queensland Department of Agriculture and Fisheries , Charters Towers, QLD 4820 , Australia
5. Livestock Production Science, KwaZulu-Natal Department of Agriculture and Rural Development , Cedara , South Africa