1. School of Nature Conservation, Beijing Forestry University, Beijing, China
2. Institute of Botany, Chinese Academy of Sciences, Beijing, China
3. College of Horticulture and Forestry Sciences, Huazhong Agricultural University, Wuhan, China
4. Institute of Wetland Ecology & Clone Ecology/Zhejiang Provincial Key Laboratory of Plant Evolutionary Ecology and Conservation, Taizhou University, Taizhou, China