1. Medical Research Council, Centre for Reproductive Health, Queen's Medical Research Institute, 47 Little France Crescent, EdinburghEH16 4TJ, UK
2. Department of Haematology/Oncology, Royal Hospital for Sick Children, 9 Sciennes Rd, Edinburgh EH9 1LF, UK
3. Institute of Cell Biology and Centre for Integrative Physiology, University of Edinburgh, Hugh Robson Building, George Square, Edinburgh EH8 9XD, UK