1. Department of Bioengineering, University of Pennsylvania , 210 S. 33rd Street, 240 Skirkanich Hall, Philadelphia, PA 19104-6321, USA
2. Department of Bioengineering, Department of Physics & Astronomy , Department of Electrical & Systems Engineering, Department of Neurology and Department of Psychiatry, University of Pennsylvania, 210 S. 33rd Street, 240 Skirkanich Hall, Philadelphia, PA 19104-6321, USA and Santa Fe Institute, Santa Fe, NM 87501, USA