1. Institute of Evolution & Marine Biodiversity, Ocean University of China, Qingdao 266003, China
2. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266003, China
3. MOE Key Laboratory of Marine Genetics and Breeding, College of Marine Life Sciences, Ocean University of China, Qingdao 266003, China
4. School of Life Science, Anhui Agricultural University, Hefei 230036, China
5. Department of Pathology, University of Michigan, Ann Arbor, MI 48109, USA